Distal leaves long-subulate beyond an elliptic, ovate or obovate base, 1.2-2.7 mm, leaf base usually strongly differentiated in shape; distal laminal cells short- to long-rectangular. Sexual condition autoicous. Seta 0.2-0.5 mm. Capsule neck short, bluntly conic. Spores reticulate. Calyptra strongly papillose.