Shrub to 4 m, the young stems glabrous to sparsely pilose, the older ones with gray exfoliating bark; petioles glabrous, 1-5 cm, avg a third as long as the blade, mostly with linear stipules; lvs round-ovate, 5-10 cm, acute, coarsely serrate with 10-20 teeth on a side, all or most deeply cordate, glabrous or sparsely hairy above, thinly hairy beneath at least on the veins; cyme 5-7-rayed, the peduncle 3-5 cm; hypanthium glandular; stylopodium glabrous; fr blue-black, flat-ellipsoid, 8-10 mm; stone flattened, slightly grooved on both sides; 2n=18, 36. Rocky woods and hills; e. Pa.; Ind. to Io., s. to Ky. and Ark. May.

Gleason, Henry A. & Cronquist, Arthur J. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. lxxv + 910 pp.

©The New York Botanical Garden. All rights reserved. Used by permission.

From Flora of Indiana (1940) by Charles C. Deam

Very local. On gravelly or rocky, wooded slopes, usually bordering streams.

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Indiana Coefficient of Conservatism: C = 10

Wetland Indicator Status: N/A

Diagnostic Traits: leaves pinnately veined, the veins +/- straight and sparingly branched, ending in marginal tooth; leaves widest at or below middle, the base cordate; petioles >5 mm; stipules present (at flowering); bud scales present; inflorescence branches from a single point.