Chamaedaphne

Shrubs. Stems erect, ascending or spreading; twigs hairy, lepidote. Leaves persistent; blade usually oblong to elliptic, rarely obovate, coriaceous, margins entire or denticulate-crenulate, plane, abaxial surface glabrous, silvery, stramineous or brownish lepidote-scaled (splitting in age); venation reticulodromous. Inflorescences terminal, (leafy) racemes, (flowers secund), 8-20-flowered, (from buds produced in previous season). Flowers: sepals 5, distinct, ovate to broadly triangular; petals 5, connate ca. 3/4 their lengths, white, corolla cylindric to urceolate-cylindric, slightly narrowed at throat, lobes much shorter than tube, (glabrous); stamens 10, included; filaments ± straight, flattened, subulate, abruptly narrowed at base, papillate, glabrous, without spurs; anther with 2 awns (awns erect, tubular, as long as body), dehiscent by terminal or subterminal pore; pistil 5-carpellate; ovary 5-locular; (style enlarged just distal to ovary); stigma slightly expanded, truncate. Fruits capsular, depressed-globose, dry-dehiscent, (with unthickened sutures). Seeds 35-50, wedge-shaped, flattened, (without wings); testa reticulate, (cells isodiametric). x = 11.

Chamaedaphne has been included in other genera of subfamily Vaccinioideae (Andromeda, Lyonia) and more recently has been placed in tribe Gaultherieae (K. A. Kron et al. 2002). Its leaf anatomy differs from that of other genera in having venation transitional between pleuroplastic and basiplastic types, denser venation, and thin vein endings (K. Lems 1964).

Fls 5-merous; cal deeply parted, persistent, closely subtended by 2 bractlets, the lobes imbricate; cor nearly cylindric, slightly narrowed at the throat; stamens 10, included; filaments flat; anthers oblong, each of the 2 pollen-sacs prolonged into an erect tube as long as the body and opening by a terminal or subterminal pore; ovary 5-locular, subtended by a 10-lobed disk; style elongate; stigma truncate; capsule depressed-globose, loculicidal; evergreen shrub with alternate lvs and numerous nodding, axillary fls forming a leafy raceme; stem and lower lf-surface and cal brown-lepidote. (Cassandra) Monospecific.

Gleason, Henry A. & Cronquist, Arthur J. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. lxxv + 910 pp.

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