Selaginella densa

Plants terrestrial or on rock, forming cushionlike or loose mats. Stems decumbent or creeping, not readily fragmenting, irregularly forked, without budlike arrested branches, tips straight; main stem upperside and underside structurally slightly different, conspicuously indeterminate, lateral branches radially symmetric, conspicuously or inconspicuously determinate, strongly ascending, 2--3-forked. Rhizophores borne on upperside of stems, throughout stem length, 0.2--0.35 mm diam. Leaves essentially monomorphic, in poorly defined pseudowhorls of 5 or 6, tightly appressed, ascending, green, linear to linear-lanceolate, (2.7--)3--5 X 0.4--0.7 mm (upperside leaves smaller than underside ones, also smaller on ascending buds); abaxial ridges present; base long-decurrent, oblique, and glabrous on underside leaves, slightly decurrent, oblique, and sometimes pubescent on upperside leaves; margins long-ciliate, cilia transparent, mostly ascending or spreading on proximal 1/2, ascending on distal 1/2, 0.07--0.17(--0.2) mm; apex slightly keeled to plane, rather obtuse, abruptly long-bristled; bristle white or transparent, puberulent, (1--)1.25--1.9 mm. Strobili solitary, (0.5--)1--3(--4) cm; sporophylls ovate-lanceolate, seldom ovate, abaxial ridges well defined, base glabrous, margins ciliate entire length or dentate near tip, apex usually long-bristled.

Prairies, alpine meadows, dry rocky slopes, rock crevices, sandstone, quartzite or granite rock, and dry gravelly, clayey or sandy soil; 1100--4000 m; Alta., B.C., Man., Ont., Sask.; Ariz., Colo., Idaho, Mont., Nebr., N.Mex., N.Dak., S.Dak., Utah, Wyo.

Selaginella densa has been treated as including three varieties: var. densa , var. scopulorum (Maxon) R. M. Tryon, and var. standleyi (Maxon) R. M. Tryon (R. M. Tryon 1955), which are recognized here at the species level. Intermediates between S . densa and the other two species of the group may represent ecological variations of the species, hybrids between species within the complex, or hybrids with other closely related species, such as S . watsonii . This group is in need of detailed systematic studies. Megasporangia with only two well-developed megaspores have been observed, which may indicate apogamy and the presence of different races as found in S . rupestris .

STEMS: monomorphic, not readily fragmenting when dry, the branch tips appearing straight when dry, prostrate, forming small, low, loose or dense cushionlike mats. RHIZOPHORES: produced throughout. LEAVES: slightly dimorphic, 2.5-5.0 mm long, (adaxial leaves sometimes slightly shorter than the abaxial leaves, linear to narrowly lanceolate, green to dark green when hydrated (adaxial leaves often persistently tan to brownish-tinged), the base decurrent (similar in color to the stem) and somewhat oblique, the tip with a straight or somewhat curved white seta 0.4-1.9 mm long (often breaking off with age), the margins ciliate. STROBILI: 1-3 mm long, the sporophylls broadly lanceolate to ovate, narrowed to an acute tip with or without a seta, the margins entirely ciliate or denticulate distally. MEGASPORES: usually relatively coarsely rugose-reticulate, orange. NOTES: 3 vars.; w U.S. e to ND and TX, AK; Can. REFERENCES: Yatskievych, G. and M. D. Windham. 2009. Vascular Plants of Arizona: Selaginellaceae. CANOTIA 5 (1): 39-48.

Common Name: lesser spikemoss Rarity: None Etymology: Selaginella is a diminutive of Selago, the name of another similar plant,