Annuals or perennials; (caudex simple or branched), [rhizomatous]; not scapose. Stems erect or ascending [decumbent], unbranched or branched distally. Leaves basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire or palmately lobed, rarely dentate or pinnatifid, (pinnately or palmately veined); cauline petiolate or sessile, blade (base cuneate or sagittate-amplexicaul), margins entire or repand [dentate, crenate], (pinnately or palmately veined, ultimate veins sometimes ending with apiculate callosities). Racemes (corymbose). Fruiting pedicels divaricate to ascending, slender or stout. Flowers: sepals ovate or oblong; petals usually spatulate, rarely obovate, claw not differentiated from blade, (apex obtuse to emarginate); stamens slightly tetradynamous; filaments slightly dilated basally; anthers ovate or oblong, (apex obtuse); nectar glands confluent, often subtending bases of stamens, median glands present or absent. Fruits sessile or shortly stipitate, linear or oblong [ovoid, lanceoloid], smooth or torulose, terete, slightly 4-angled, latiseptate, or angustiseptate; valves each with obscure or prominent midvein; replum rounded; septum complete or perforated; ovules [2-] (6-)8-96 per ovary; stigma capitate. Seeds usually uniseriate, rarely biseriate, plump, not winged, oblong or ovoid; seed coat (usually obscurely reticulate, rarely foveolate or papillate), often not mucilaginous when wetted; cotyledons incumbent [rarely accumbent].
This project was made possible in part by the Institute of Museum and Library Services [MG-70-19-0057-19].