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           [Tracheobionta]
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             [Angiospermae]
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             Core Eudicots
               Rosids
               Fabids
                    Cucurbitales
                            <b>Begoniaceae</b>
                                     <i>Begonia</i>
                                           [Lophophytum]
                                             <i>Begonia abbottii</i>
                                             <i>Begonia aborensis</i>
                                             <i>Begonia acclivis</i>
                                             <i>Begonia acerifolia</i>
                                             <i>Begonia aconitifolia</i>
                                                   [Begonia sceptrum]
                                             <i>Begonia acuminatissima</i>
                                             <i>Begonia acutifolia</i>
                                             <i>Begonia acutiloba</i>
                                             <i>Begonia adpressa</i>
                                             <i>Begonia aequata</i>
                                             <i>Begonia aequatoguineensis</i>
                                             <i>Begonia aggeloptera</i>
                                             <i>Begonia agusanensis</i>
                                             <i>Begonia almedana</i>
                                             <i>Begonia alnifolia</i>
                                             <i>Begonia amoeboides</i>
                                             <i>Begonia ampla</i>
                                                   [Begonia duruensis]
                                             <i>Begonia androturba</i>
                                             <i>Begonia angolensis</i>
                                             <i>Begonia angulata</i>
                                             <i>Begonia angustiloba</i>
                                             <i>Begonia anisoptera</i>
                                             <i>Begonia anisosepala</i>
                                                   [Begonia calophylla]
                                             <i>Begonia annobonensis</i>
                                                   [Begonia quintasii]
                                             <i>Begonia antongilensis</i>
                                                   [Begonia antongilensis var. cuneata]
                                             <i>Begonia aptera</i>
                                             <i>Begonia arborescens</i>
                                             <i>Begonia asperifolia</i>
                                             <i>Begonia aspleniifolia</i>
                                                   [Begonia filicifolia]
                                             <i>Begonia assurgens</i>
                                             <i>Begonia asteroides</i>
                                             <i>Begonia atroglandulosa</i>
                                               <i>Begonia atroglandulosa subsp. atroglandulosa</i>
                                               <i>Begonia atroglandulosa subsp. tshelaensis</i>
                                             <i>Begonia augustae</i>
                                             <i>Begonia auriculata</i>
                                                   [Begonia sciaphila var. longipedunculata]
                                             <i>Begonia baccata</i>
                                                   [Begonia crateris]
                                             <i>Begonia balansae</i>
                                             <i>Begonia balmisiana</i>
                                                   [Begonia balmisiana var. mitelliflora]
                                                   [Begonia balmisiana var. mitellifolia]
                                                   [Begonia monoptera]
                                             <i>Begonia bangii</i>
                                             <i>Begonia barahonensis</i>
                                                   [Begonia plumieri var. barahonensis]
                                             <i>Begonia barkeri</i>
                                                   [Begonia megaphylla]
                                             <i>Begonia bartlettiana</i>
                                             <i>Begonia beccariana</i>
                                             <i>Begonia bequaertii</i>
                                             <i>Begonia beryllae</i>
                                             <i>Begonia bicolor</i>
                                             <i>Begonia bipinnatifida</i>
                                             <i>Begonia biserrata</i>
                                                   [Begonia crenatiflora]
                                                   [Begonia palmaris var. jurgensenii]
                                                   [Begonia palmaris]
                                               <i>Begonia biserrata var. glandulosa</i>
                                             <i>Begonia boissieri</i>
                                                   [Begonia abaculoides]
                                             <i>Begonia boliviensis</i>
                                             <i>Begonia bolleana</i>
                                             <i>Begonia bonus-henricus</i>
                                             <i>Begonia borneensis</i>
                                             <i>Begonia bosseri</i>
                                             <i>Begonia brachybotrys</i>
                                                   [Begonia brachyptera]
                                             <i>Begonia brachypoda</i>
                                                   [Begonia brachypoda var. pilosula]
                                             <i>Begonia bracteata</i>
                                             <i>Begonia bracteosa</i>
                                             <i>Begonia breedlovei</i>
                                             <i>Begonia brevibracteata</i>
                                             <i>Begonia brevipes</i>
                                             <i>Begonia brevirimosa</i>
                                             <i>Begonia burbidgei</i>
                                             <i>Begonia burkillii</i>
                                             <i>Begonia buseyi</i>
                                             <i>Begonia calderonii</i>
                                                   [Begonia falcata]
                                             <i>Begonia californica</i>
                                             <i>Begonia camiguinensis</i>
                                             <i>Begonia campanensis</i>
                                             <i>Begonia capillipes</i>
                                                   [Begonia cultrata]
                                             <i>Begonia cardiocarpa</i>
                                                   [Begonia bakeri]
                                             <i>Begonia carletonii</i>
                                             <i>Begonia carolineifolia</i>
                                             <i>Begonia caroliniifolia</i>
                                             <i>Begonia carpinifolia</i>
                                             <i>Begonia carrieae</i>
                                             <i>Begonia cavallyensis</i>
                                             <i>Begonia cebadillensis</i>
                                             <i>Begonia chaetocarpa</i>
                                                   [Begonia chaetocarpa var. glabriflora]
                                             <i>Begonia chiapensis</i>
                                             <i>Begonia chingipengii</i>
                                             <i>Begonia chitoensis</i>
                                             <i>Begonia chlorolepis</i>
                                             <i>Begonia ciliifera</i>
                                             <i>Begonia ciliobracteata</i>
                                                   [Begonia cameroonensis]
                                                   [Begonia dielsiana]
                                                   [Begonia hookeriana]
                                                   [Begonia raynaliorum]
                                             <i>Begonia clemensiae</i>
                                             <i>Begonia clypeifolia</i>
                                               <i>Begonia clypeifolia subsp. celer</i>
                                               <i>Begonia clypeifolia subsp. clypeifolia</i>
                                               <i>Begonia clypeifolia subsp. mayombensis</i>
                                             <i>Begonia coccinea</i>
                                             <i>Begonia conchifolia</i>
                                             <i>Begonia confusa</i>
                                             <i>Begonia conipila</i>
                                             <i>Begonia contracta</i>
                                             <i>Begonia convallariodora</i>
                                             <i>Begonia convolvulacea</i>
                                             <i>Begonia cooperi</i>
                                             <i>Begonia copeyana</i>
                                             <i>Begonia cordifolia</i>
                                             <i>Begonia coronensis</i>
                                             <i>Begonia corredorana</i>
                                             <i>Begonia crassicaulis</i>
                                             <i>Begonia crenata</i>
                                             <i>Begonia crispipila</i>
                                             <i>Begonia cristobalensis</i>
                                             <i>Begonia croatii</i>
                                             <i>Begonia cryptocarpa</i>
                                             <i>Begonia cuatrecasasiana</i>
                                             <i>Begonia cucullata</i>
                                               <i>Begonia cucullata var. arenosicola</i>
                                                     [Begonia subcucullata]
                                               <i>Begonia cucullata var. hookeri</i>
                                               <i>Begonia cucullata var. spatulata</i>
                                             <i>Begonia culasiensis</i>
                                             <i>Begonia cumingiana</i>
                                             <i>Begonia cumingii</i>
                                                   [Begonia philippinensis]
                                             <i>Begonia cylindrata</i>
                                             <i>Begonia dealbata</i>
                                             <i>Begonia decandra</i>
                                             <i>Begonia decora</i>
                                             <i>Begonia dewildei</i>
                                             <i>Begonia dioica</i>
                                                   [Begonia amoena]
                                             <i>Begonia dipetala</i>
                                             <i>Begonia dodsonii</i>
                                             <i>Begonia domingensis</i>
                                             <i>Begonia dregei</i>
                                             <i>Begonia duncan-thomasii</i>
                                             <i>Begonia dusenii</i>
                                             <i>Begonia ebolowensis</i>
                                                   [Begonia lethomasiae]
                                                   [Begonia sanjeensis]
                                             <i>Begonia echinosepala</i>
                                             <i>Begonia edulis</i>
                                             <i>Begonia egregia</i>
                                             <i>Begonia elaeagnifolia</i>
                                                   [Begonia schultzei]
                                                   [Begonia wilczekiana]
                                             <i>Begonia elatostemmoides</i>
                                                   [Begonia loloensis]
                                             <i>Begonia elegans</i>
                                             <i>Begonia elnidoensis</i>
                                             <i>Begonia eminii</i>
                                                   [Begonia ealensis]
                                                   [Begonia macrostyla]
                                                   [Begonia poggei var. albiflora]
                                                   [Begonia poggei]
                                                   [Begonia warburgii]
                                             <i>Begonia engleri</i>
                                                   [Begonia engleri var. nuda]
                                             <i>Begonia epipsila</i>
                                             <i>Begonia erectocaulis</i>
                                             <i>Begonia erectotricha</i>
                                             <i>Begonia erythrocarpa</i>
                                                   [Begonia pennellii]
                                             <i>Begonia esculenta</i>
                                             <i>Begonia estrellensis</i>
                                             <i>Begonia exilis</i>
                                             <i>Begonia extranea</i>
                                             <i>Begonia falcifolia</i>
                                             <i>Begonia falciloba</i>
                                             <i>Begonia faustinoi</i>
                                             <i>Begonia fenicis</i>
                                             <i>Begonia fernaldiana</i>
                                             <i>Begonia ferramica</i>
                                             <i>Begonia ferruginea</i>
                                                   [Begonia ferruginea var. dilatata]
                                             <i>Begonia fimbriata</i>
                                             <i>Begonia fimbristipula</i>
                                             <i>Begonia fischeri</i>
                                                   [Begonia fischeri var. tovarensis]
                                                   [Begonia tovarensis]
                                             <i>Begonia foliosa</i>
                                                   [Begonia microphylla]
                                               <i>Begonia foliosa var. foliosa</i>
                                               <i>Begonia foliosa var. miniata</i>
                                             <i>Begonia forgetiana</i>
                                             <i>Begonia formosana</i>
                                             <i>Begonia foxworthyi</i>
                                             <i>Begonia fruticosa</i>
                                             <i>Begonia fuchsioides</i>
                                                   [Begonia fuchsioides var. miniata]
                                             <i>Begonia furfuracea</i>
                                             <i>Begonia fusca</i>
                                             <i>Begonia fusialata</i>
                                               <i>Begonia fusialata var. fusialata</i>
                                                     [Begonia alepensis]
                                               <i>Begonia fusialata var. parviflora</i>
                                               <i>Begonia fusialata var. principensis</i>
                                             <i>Begonia fusibulba</i>
                                             <i>Begonia fusicarpa</i>
                                             <i>Begonia gabonensis</i>
                                             <i>Begonia gentilii</i>
                                             <i>Begonia georgei</i>
                                             <i>Begonia gironellae</i>
                                             <i>Begonia gitingensis</i>
                                             <i>Begonia glabra</i>
                                                   [Begonia glabra var. amplifolia]
                                                   [Begonia locellata]
                                                   [Begonia physalifolia]
                                             <i>Begonia glandulifera</i>
                                             <i>Begonia glandulosa</i>
                                                   [Begonia hidalgensis]
                                             <i>Begonia glaucophylla</i>
                                             <i>Begonia goegoensis</i>
                                             <i>Begonia gossweileri</i>
                                             <i>Begonia goudotii</i>
                                             <i>Begonia gracilior</i>
                                             <i>Begonia gracilipes</i>
                                             <i>Begonia gracilis</i>
                                               <i>Begonia gracilis var. martiana</i>
                                               <i>Begonia gracilis var. nervipilosa</i>
                                             <i>Begonia grandis</i>
                                                   [Begonia evansiana]
                                               <i>Begonia grandis subsp. sinensis</i>
                                             <i>Begonia griffithiana</i>
                                             <i>Begonia grisea</i>
                                             <i>Begonia guaduensis</i>
                                             <i>Begonia gueritziana</i>
                                             <i>Begonia gutierrezii</i>
                                             <i>Begonia haageana</i>
                                             <i>Begonia halconensis</i>
                                             <i>Begonia hatacoa</i>
                                             <i>Begonia hemicardia</i>
                                             <i>Begonia heracleifolia</i>
                                                   [Begonia heracleifolia var. nigricans]
                                             <i>Begonia heterochroma</i>
                                             <i>Begonia heydei</i>
                                                   [Begonia triloba]
                                             <i>Begonia hintoniana</i>
                                             <i>Begonia hirsuta</i>
                                                   [Begonia filipes]
                                                   [Begonia heterodonta]
                                                   [Begonia mameiana]
                                             <i>Begonia hirsutula</i>
                                                   [Begonia comperei]
                                                   [Begonia klainei]
                                                   [Begonia subfalcata]
                                             <i>Begonia hirta</i>
                                               <i>Begonia hirta var. cordifolia</i>
                                             <i>Begonia hirtella</i>
                                             <i>Begonia hitchcockii</i>
                                             <i>Begonia holtonis</i>
                                             <i>Begonia hondurensis</i>
                                             <i>Begonia horticola</i>
                                                   [Begonia parva]
                                                   [Begonia spraguei]
                                             <i>Begonia humilis</i>
                                             <i>Begonia hydrocotylifolia</i>
                                                   [Begonia asarifolia]
                                             <i>Begonia ignea</i>
                                             <i>Begonia imperialis</i>
                                             <i>Begonia incana</i>
                                             <i>Begonia incarnata</i>
                                                   [Begonia metallica]
                                             <i>Begonia incisa</i>
                                             <i>Begonia incisoserrata</i>
                                             <i>Begonia integrifolia</i>
                                             <i>Begonia involucrata</i>
                                                   [Begonia broussonetiifolia]
                                                   [Begonia laciniosa]
                                             <i>Begonia isabelensis</i>
                                             <i>Begonia isoptera</i>
                                                   [Begonia angustifolia]
                                                   [Begonia geniculata]
                                             <i>Begonia itapavensis</i>
                                             <i>Begonia itupavensis</i>
                                             <i>Begonia iucunda</i>
                                             <i>Begonia jagorii</i>
                                             <i>Begonia jaliscana</i>
                                             <i>Begonia jamaicensis</i>
                                             <i>Begonia johnstonii</i>
                                                   [Begonia johnstonii f. pilosa]
                                             <i>Begonia josephi</i>
                                             <i>Begonia juntasensis</i>
                                             <i>Begonia kalbreyeri</i>
                                               <i>Begonia kalbreyeri var. orquidensis</i>
                                             <i>Begonia karperi</i>
                                             <i>Begonia karwinskyana</i>
                                             <i>Begonia kenworthyae</i>
                                             <i>Begonia killipiana</i>
                                             <i>Begonia kisuluana</i>
                                                   [Begonia zobiaensis]
                                             <i>Begonia komoensis</i>
                                             <i>Begonia labordei</i>
                                             <i>Begonia lachaoensis</i>
                                             <i>Begonia lacunosa</i>
                                                   [Begonia mayombensis]
                                             <i>Begonia lagunensis</i>
                                             <i>Begonia laminariae</i>
                                             <i>Begonia lanceolata</i>
                                             <i>Begonia langbianensis</i>
                                             <i>Begonia laporteifolia</i>
                                                   [Begonia hypogaea]
                                                   [Begonia schlechteri]
                                             <i>Begonia latistipula</i>
                                             <i>Begonia leptantha</i>
                                             <i>Begonia letestui</i>
                                             <i>Begonia letouzeyi</i>
                                             <i>Begonia leuconeura</i>
                                             <i>Begonia lignescens</i>
                                             <i>Begonia lindleyana</i>
                                             <i>Begonia listada</i>
                                             <i>Begonia loheri</i>
                                             <i>Begonia longibractea</i>
                                             <i>Begonia longifolia</i>
                                             <i>Begonia longipetiolata</i>
                                                   [Begonia bipindensis]
                                                   [Begonia crassipes]
                                                   [Begonia gladiifolia]
                                                   [Begonia gracilipetiolata]
                                                   [Begonia kribensis]
                                                   [Begonia ludwigsii]
                                                   [Begonia macrura]
                                                   [Begonia nicolai-hallei]
                                                   [Begonia squamulosa var. bipindensis]
                                             <i>Begonia longirostris</i>
                                             <i>Begonia longitepala</i>
                                             <i>Begonia lopensis</i>
                                             <i>Begonia lophoptera</i>
                                             <i>Begonia loranthoides</i>
                                               <i>Begonia loranthoides subsp. loranthoides</i>
                                                     [Begonia henriquesii]
                                               <i>Begonia loranthoides subsp. rhopalocarpa</i>
                                                     [Begonia injoloensis]
                                                     [Begonia rhopalocarpa]
                                             <i>Begonia louis-williamsii</i>
                                             <i>Begonia ludicra</i>
                                                   [Begonia liebmannii]
                                             <i>Begonia lunaris</i>
                                             <i>Begonia luxurians</i>
                                             <i>Begonia luzonensis</i>
                                             <i>Begonia lyman-smithii</i>
                                             <i>Begonia lyniceorum</i>
                                             <i>Begonia macrocarpa</i>
                                                   [Begonia bruneelii]
                                                   [Begonia gouroana]
                                                   [Begonia gracilicaulis]
                                                   [Begonia pseudimpatiens]
                                                   [Begonia romeensis]
                                                   [Begonia simii]
                                             <i>Begonia maculata</i>
                                             <i>Begonia macvaughii</i>
                                             <i>Begonia malabarica</i>
                                             <i>Begonia malindangensis</i>
                                             <i>Begonia manicata</i>
                                                   [Begonia manicata var. manicativiridi]
                                                   [Begonia manicata var. ocozocoautlaensis]
                                                   [Begonia manicata var. peltata]
                                                   [Begonia robustior]
                                             <i>Begonia manillensis</i>
                                             <i>Begonia mannii</i>
                                                   [Begonia epiphytica]
                                                   [Begonia excelsa]
                                             <i>Begonia mariti</i>
                                             <i>Begonia martiana</i>
                                             <i>Begonia masoniana</i>
                                             <i>Begonia maxima</i>
                                             <i>Begonia maynensis</i>
                                             <i>Begonia mazae</i>
                                             <i>Begonia mbangaensis</i>
                                             <i>Begonia mearnsii</i>
                                             <i>Begonia media</i>
                                             <i>Begonia megalantha</i>
                                             <i>Begonia merrittii</i>
                                             <i>Begonia meyeri-johannis</i>
                                             <i>Begonia michoacana</i>
                                             <i>Begonia micranthera</i>
                                             <i>Begonia microsperma</i>
                                                   [Begonia ficicola]
                                             <i>Begonia mildbraedii</i>
                                             <i>Begonia militaris</i>
                                             <i>Begonia mindanaensis</i>
                                             <i>Begonia mindorensis</i>
                                                   [Begonia sordidissima]
                                             <i>Begonia minicarpa</i>
                                                   [Begonia modesta]
                                             <i>Begonia minor</i>
                                                   [Begonia nitida]
                                             <i>Begonia minuta</i>
                                             <i>Begonia minutifolia</i>
                                             <i>Begonia molinana</i>
                                             <i>Begonia molleri</i>
                                                   [Mezierea molleri]
                                             <i>Begonia monadelpha</i>
                                             <i>Begonia monophylla</i>
                                             <i>Begonia monte-alenensis</i>
                                             <i>Begonia monticola</i>
                                             <i>Begonia montis-elephantis</i>
                                             <i>Begonia motozintlensis</i>
                                             <i>Begonia mucronistipula</i>
                                                   [Begonia davidsoniae]
                                             <i>Begonia multangula</i>
                                             <i>Begonia multinervia</i>
                                                   [Begonia cuspidata]
                                             <i>Begonia multistaminea</i>
                                             <i>Begonia ndongensis</i>
                                             <i>Begonia negrosensis</i>
                                             <i>Begonia nelumbiifolia</i>
                                             <i>Begonia nelumbonifolia</i>
                                             <i>Begonia nemoralis</i>
                                             <i>Begonia neopurpurea</i>
                                                   [Begonia purpurea]
                                             <i>Begonia nepalensis</i>
                                                   [Begonia gigantea]
                                             <i>Begonia nigritarum</i>
                                                   [Begonia merrillii]
                                             <i>Begonia nurii</i>
                                             <i>Begonia nyassensis</i>
                                             <i>Begonia oaxacana</i>
                                                   [Begonia luxii]
                                                   [Begonia oaxacana var. pilosula]
                                             <i>Begonia obliqua</i>
                                                   [Begonia martinicensis]
                                                   [Begonia odorata]
                                             <i>Begonia octopetala</i>
                                             <i>Begonia odeteiantha</i>
                                             <i>Begonia ornithocarpa</i>
                                             <i>Begonia oxyanthera</i>
                                                   [Begonia jussiaeicarpa]
                                             <i>Begonia oxyloba</i>
                                                   [Begonia cladocarpa]
                                                   [Begonia conraui]
                                                   [Begonia heddei]
                                                   [Begonia kummeriae]
                                                   [Begonia lehmbachii]
                                                   [Begonia petrophila]
                                                   [Begonia pycnocaulis]
                                                   [Begonia sassandrensis]
                                                   [Begonia seretii]
                                                   [Begonia togoensis]
                                             <i>Begonia oxysperma</i>
                                             <i>Begonia palawanensis</i>
                                             <i>Begonia paleata</i>
                                             <i>Begonia palmata</i>
                                                   [Begonia laciniata]
                                               <i>Begonia palmata var. khasiana</i>
                                               <i>Begonia palmata var. palmata</i>
                                             <i>Begonia palmeri</i>
                                                   [Begonia macdanielsii]
                                             <i>Begonia panayensis</i>
                                             <i>Begonia papuana</i>
                                             <i>Begonia parviflora</i>
                                                   [Begonia myriantha]
                                             <i>Begonia parvilimba</i>
                                             <i>Begonia pavonina</i>
                                             <i>Begonia pedata</i>
                                             <i>Begonia pedatifida</i>
                                             <i>Begonia pedunculosa</i>
                                             <i>Begonia pelargoniiflora</i>
                                             <i>Begonia peltata</i>
                                             <i>Begonia peperomioides</i>
                                             <i>Begonia pernambucensis</i>
                                             <i>Begonia peruviana</i>
                                             <i>Begonia picta</i>
                                             <i>Begonia pilosella</i>
                                             <i>Begonia pinetorum</i>
                                             <i>Begonia pinnatifida</i>
                                             <i>Begonia piurensis</i>
                                             <i>Begonia plantaginea</i>
                                             <i>Begonia platyptera</i>
                                             <i>Begonia plebeja</i>
                                             <i>Begonia pleiopetala</i>
                                             <i>Begonia plumieri</i>
                                             <i>Begonia pluvialis</i>
                                             <i>Begonia poculifera</i>
                                               <i>Begonia poculifera var. poculifera</i>
                                                     [Begonia adolfi-friderici]
                                                     [Begonia haullevilleana]
                                               <i>Begonia poculifera var. teusziana</i>
                                                     [Begonia teusziana]
                                             <i>Begonia polyandra</i>
                                             <i>Begonia polygonata</i>
                                                   [Begonia fonsecae]
                                             <i>Begonia polygonoides</i>
                                                   [Begonia cataractarum]
                                                   [Begonia epilobioides]
                                                   [Begonia rhipsaloides]
                                             <i>Begonia popenoei</i>
                                             <i>Begonia porteana</i>
                                             <i>Begonia portillana</i>
                                             <i>Begonia potamophila</i>
                                                   [Begonia batesii]
                                                   [Begonia macropoda]
                                             <i>Begonia preussii</i>
                                                   [Begonia buchholzii]
                                                   [Begonia sessilanthera]
                                             <i>Begonia princeae</i>
                                                   [Begonia homblei]
                                                   [Begonia princeae f. grossidentata]
                                                   [Begonia princeae f. racemigera]
                                                   [Begonia princeae f. rhodesiana]
                                                   [Begonia princeae f. vulgata]
                                                   [Begonia princeae var. racemigera]
                                                   [Begonia subacuto-alata]
                                                   [Begonia verdickii]
                                                   [Begonia wellmanii]
                                             <i>Begonia pringlei</i>
                                             <i>Begonia prismatocarpa</i>
                                               <i>Begonia prismatocarpa subsp. delobata</i>
                                               <i>Begonia prismatocarpa subsp. petraea</i>
                                                     [Begonia petraea]
                                               <i>Begonia prismatocarpa subsp. prismatocarpa</i>
                                             <i>Begonia pryeriana</i>
                                             <i>Begonia pseudolateralis</i>
                                             <i>Begonia pseudoviola</i>
                                                   [Begonia subtilis]
                                             <i>Begonia puberula</i>
                                             <i>Begonia pubescens</i>
                                             <i>Begonia pudica</i>
                                             <i>Begonia pulchella</i>
                                             <i>Begonia pulcherrima</i>
                                             <i>Begonia pumilio</i>
                                             <i>Begonia purpusii</i>
                                             <i>Begonia pustulata</i>
                                             <i>Begonia pycnantha</i>
                                             <i>Begonia pygmaea</i>
                                             <i>Begonia quadrialata</i>
                                               <i>Begonia quadrialata subsp. dusenii</i>
                                               <i>Begonia quadrialata subsp. nimbaensis</i>
                                                     [Begonia leichtiana]
                                               <i>Begonia quadrialata subsp. quadrialata</i>
                                               <i>Begonia quadrialata var. pilosa</i>
                                               <i>Begonia quadrialata var. quadrialata</i>
                                                     [Begonia calabarica]
                                                     [Begonia modica]
                                                     [Begonia poikilantha]
                                                     [Begonia quadrialata var. speciosa]
                                                     [Begonia whytei]
                                             <i>Begonia quaternata</i>
                                             <i>Begonia quercifolia</i>
                                             <i>Begonia radicans</i>
                                                   [Begonia procumbens]
                                             <i>Begonia ramosii</i>
                                             <i>Begonia relicta</i>
                                             <i>Begonia reniformis</i>
                                             <i>Begonia rex</i>
                                             <i>Begonia rhodochlamys</i>
                                             <i>Begonia ricinifolia</i>
                                             <i>Begonia riparia</i>
                                             <i>Begonia rosea</i>
                                             <i>Begonia rossmanniae</i>
                                             <i>Begonia rostrata</i>
                                               <i>Begonia rostrata var. argutiserrata</i>
                                               <i>Begonia rostrata var. brachyptera</i>
                                               <i>Begonia rostrata var. rostrata</i>
                                                     [Begonia chevalieri]
                                                     [Begonia elliotii]
                                             <i>Begonia rubella</i>
                                             <i>Begonia rubrifolia</i>
                                             <i>Begonia rubromarginata</i>
                                             <i>Begonia rubronervata</i>
                                             <i>Begonia rufa</i>
                                                   [Begonia lobata]
                                             <i>Begonia rumpiensis</i>
                                             <i>Begonia rwandensis</i>
                                             <i>Begonia salisburyana</i>
                                             <i>Begonia samarensis</i>
                                             <i>Begonia sandtii</i>
                                             <i>Begonia sanguinea</i>
                                             <i>Begonia santarosensis</i>
                                             <i>Begonia sartorii</i>
                                                   [Begonia lobulata]
                                                   [Begonia sarcophylla]
                                             <i>Begonia saxicola</i>
                                             <i>Begonia scapigera</i>
                                               <i>Begonia scapigera subsp. australis</i>
                                               <i>Begonia scapigera subsp. scapigera</i>
                                             <i>Begonia schaeferi</i>
                                                   [Begonia ramosa]
                                             <i>Begonia scharffii</i>
                                                   [Begonia scharffiana]
                                             <i>Begonia schliebenii</i>
                                             <i>Begonia schulziana</i>
                                             <i>Begonia sciaphila</i>
                                             <i>Begonia scutifolia</i>
                                                   [Begonia triflora var. caloskiadia]
                                                   [Begonia triflora]
                                             <i>Begonia scutulum</i>
                                             <i>Begonia seemanniana</i>
                                                   [Begonia seemanniana var. longistipulacea]
                                             <i>Begonia semiovata</i>
                                                   [Begonia flexuosa]
                                                   [Begonia guyanensis var. glaberrima]
                                                   [Begonia guyanensis]
                                                   [Begonia spruceana]
                                             <i>Begonia semperflorens</i>
                                             <i>Begonia sericoneura</i>
                                                   [Begonia nicaraguensis]
                                                   [Begonia pilifera]
                                             <i>Begonia sessilifolia</i>
                                                   [Begonia gilgii]
                                             <i>Begonia sikkimensis</i>
                                               <i>Begonia sikkimensis var. kamengensis</i>
                                             <i>Begonia silletensis</i>
                                             <i>Begonia sinensis</i>
                                             <i>Begonia sinuata</i>
                                             <i>Begonia soli-mutata</i>
                                             <i>Begonia sonderiana</i>
                                                   [Begonia sonderiana var. transgrediens]
                                             <i>Begonia sorsogonensis</i>
                                             <i>Begonia sosefiana</i>
                                             <i>Begonia sousae</i>
                                             <i>Begonia spathulata</i>
                                             <i>Begonia squamulosa</i>
                                             <i>Begonia squarrosa</i>
                                             <i>Begonia staudtii</i>
                                                   [Begonia staudtii var. dispersipilosa]
                                             <i>Begonia stellata</i>
                                             <i>Begonia stenotepala</i>
                                             <i>Begonia stigmosa</i>
                                                   [Begonia mexiae]
                                             <i>Begonia stolzii</i>
                                             <i>Begonia strigillosa</i>
                                                   [Begonia tinctoria]
                                             <i>Begonia subalpestris</i>
                                                   [Begonia macambrarensis]
                                             <i>Begonia subciliata</i>
                                               <i>Begonia subciliata var. persicina</i>
                                             <i>Begonia subcostata</i>
                                             <i>Begonia suborbiculata</i>
                                             <i>Begonia subscutata</i>
                                                   [Begonia latipetiolata]
                                             <i>Begonia subvillosa</i>
                                                   [Begonia leptotricha]
                                                   [Begonia schmidtiana]
                                             <i>Begonia susaniae</i>
                                             <i>Begonia sutherlandii</i>
                                               <i>Begonia sutherlandii subsp. latior</i>
                                                     [Begonia flava]
                                                     [Begonia sutherlandii f. densiserrata]
                                                     [Begonia sutherlandii var. latior]
                                               <i>Begonia sutherlandii subsp. sutherlandii</i>
                                                     [Begonia gueinziana]
                                             <i>Begonia sykakiengii</i>
                                             <i>Begonia tafiensis</i>
                                             <i>Begonia taiwaniana</i>
                                             <i>Begonia tapatia</i>
                                             <i>Begonia tatoniana</i>
                                                   [Begonia eminii subsp. ambacensis]
                                             <i>Begonia tawaensis</i>
                                             <i>Begonia tayloriana</i>
                                             <i>Begonia thaipingensis</i>
                                             <i>Begonia thiemei</i>
                                             <i>Begonia thomeana</i>
                                             <i>Begonia thomsonii</i>
                                             <i>Begonia tiliifolia</i>
                                             <i>Begonia tlapensis</i>
                                             <i>Begonia tonduzii</i>
                                                   [Begonia allenii]
                                             <i>Begonia trapa</i>
                                             <i>Begonia trichosepala</i>
                                                   [Begonia erythrotricha]
                                             <i>Begonia tuberosa</i>
                                             <i>Begonia turrialbae</i>
                                             <i>Begonia udisilvestris</i>
                                             <i>Begonia ulmifolia</i>
                                             <i>Begonia umbratica</i>
                                             <i>Begonia uniflora</i>
                                             <i>Begonia unifolia</i>
                                             <i>Begonia urophylla</i>
                                                   [Begonia santae-martae]
                                             <i>Begonia urticae</i>
                                                   [Begonia antioquensis]
                                                   [Begonia chiriquensis]
                                                   [Begonia urticae var. hispida]
                                             <i>Begonia uruapensis</i>
                                                   [Begonia uruapensis var. rosei]
                                             <i>Begonia valida</i>
                                             <i>Begonia vankerckhovenii</i>
                                             <i>Begonia veitchii</i>
                                             <i>Begonia venosa</i>
                                             <i>Begonia vestita</i>
                                             <i>Begonia villipetiola</i>
                                             <i>Begonia vincentiana</i>
                                             <i>Begonia violifolia</i>
                                             <i>Begonia viscida</i>
                                             <i>Begonia vitifolia</i>
                                             <i>Begonia vittariifolia</i>
                                             <i>Begonia wakefieldii</i>
                                                   [Begonia wakefieldii f. dentatiloba]
                                             <i>Begonia wallichiana</i>
                                                   [Begonia franconis]
                                             <i>Begonia weberlingii</i>
                                             <i>Begonia wilksii</i>
                                             <i>Begonia wollastonii</i>
                                                   [Begonia abyssinica]
                                                   [Begonia keniensis]
                                                   [Begonia lebrunii]
                                             <i>Begonia wrayi</i>
                                             <i>Begonia wrightiana</i>
                                             <i>Begonia x tuberhybrida</i>
                                                   [Begonia x tuberosa]
                                             <i>Begonia xanthina</i>
                                             <i>Begonia xilitlensis</i>
                                             <i>Begonia yunckeri</i>
                                                   [Begonia salvadorensis]
                                             <i>Begonia zairensis</i>
                                               <i>Begonia zairensis var. montana</i>
                                               <i>Begonia zairensis var. zairensis</i>
                                             <i>Begonia zenkeriana</i>
                                                   [Begonia zenkeri]
                                             <i>Begonia zimmermannii</i>
                                             <i>Begonia × erythrophylla</i>
                                             <i>Begonia × feastii</i>
                                             <i>Begonia × hybrida</i>
                                             <i>Begonia × tuberhybrida</i>
                                     <i>Hillebrandia</i>
                                             <i>Hillebrandia sandwicensis</i>
                                     <i>Lepsia</i>
                                     <i>Mezierea</i>